instead of a star, there's a sphere in the sky that eternally issues forth sound.
there's no light, but the sounding coming down and bouncing is analogous to sunlight. animals evolve to use a sort passive echolocation that works better during the "day".
prey animals tend to have ears that point outward, to either side, so they can have an almost 360 degree view of any predators who might try to approach. predators tend to have ears moved further forward on the head, which face forward and enable binocular vision.
a species much like giant jumping spiders evolves. instead of two main eyes, they have two main front ears. they also have a number of smaller side ears. they use hydraulic legs to leap at prey from a distance.
they evolve mandibles dextrous enough to make and use tools. at this very moment, one uses a hearing trumpet to listen to the sky, wondering if anyone else is out there.